Empires Of the Undergrowth All Species of Ants

Welcome, guideoui.com visitors. In this guide, We try to focus on Empires Of the Undergrowth All Species of Ants. It will talk about all of the species from the starting ants to the most recently added ones. This guide will be improved upon as the species of the games are introduced or updated or even become playable.
While writing this guide, we pick up many pieces of information from several sites for you. We hope that this guide will help you.

The Empire of the Undergrowth Species Guide talks about ‘all’ of the species of Ants, both Playable and Non currently available in the game, to help you learn about what each one does and strategize with and against them.

All Species of Ants

Formica Ereptor – Gene Thieves (Player Ant)

Formica Ereptor, the Gene Thief Ant Species, not a real ant species and only exists in the game, is the player’s controlled ant species. The Queen of the Species at first starts out rather pathetic compared to what it’s name suggests, only capable of producing a single type of ant at first, the worker.

However, this is only at first, as every ant colony starts out similiarly to what the player starts with, a single Queen and a few workers.

Where Ereptor’s strengths come into play is the fact that, unlike the other species of ants, Formica Ereptor can use the genetic strains of ‘other’ species, and build their ants as if they were one and the same species.

This makes Ereptor the ultimate ant species, as the others cannot do this and are stuck with their own species which will be discussed in their own seperate sections of the guide.

Formica Ereptor Queen

Formica Ereptor’s queen is like all other Queens, a giant ant that sits in the center of your colony and does nothing by herself…

She cannot attack, she cannot defend herself, but unlike all the other ant queens, she possesses unique abilities, and can activate two of four powers when her health drops below 75%

She can summon the Royal Guard, a squad of 6-7 giant Ereptor ants that have high health and strong attacks around her that will attack and defend the queen from attackers to the death, for a set period of time.

She can unleash a hoard of ants for free for a set period of time, allowing nursery tiles to be refilled with eggs even if empty and hatch their ants wihtout needing to pay for them instantly…

She can stun enemies around her and in the whole nest for a period of time

Or she can restore her health and fully plant and release a new group of ants from their nursery tiles for free

Formica Ereptor Worker

The first and Only ant available to Ereptor at the start, the ereptor worker fullfills two jobs crucial to the colony’s survival and thriving.

While any ant can collect food, only the workers can either Dig tunnels or chambers, plant down nursery tiels, plant down food storage tiles, plant down highway tiles, or pick up and plant larvae from the queen and to their assigned nursery tiles…

They are needed and despite their small size, low health, and weak attacks, can defend themselves if their numbers are high enough. However, workers should only be used for attacking or defending as a last resort, as they much prefer to do their duties as egg carriers to the queen while the soldiers do the real fighting…

All other species of ants that Ereptor can use are covered later on, and will have a (Ereptor) word next to the ant type’s name…

Formica Fusca – Black Ant’s Basics

Formica Fusca is the first ant introduced to us in the game, and while Formica Fusca has very little diversity, only two ant types to the colony, it fullfills the purpose in the Ant universe and the game, to the tutorials and basics in the game and the Undergrowth as an empire…

Formica Fusca’s Queen is just a giant fat ant that sits there and does nothing, she lays eggs and if she dies the colony dies

As for the two Ants available to Fusca… they are Worker and Soldier

Formica Fusca Worker Ant

Formica Fusca’s worker is a basic ant, it performs all the functions of the workers, similiar to Ereptor’s worker. It can gather food like other ants, and picks up eggs from the queen, delivers them to nursery tiles that the player builds, places down food storage and highway tiles.

Unlike Ereptor’s Workers though, Fusca workers tend to be also capable soldiers, as i’ve personally noticed that workers can deal slightly more damage and are best used in large numbers against single enemy targets. Don’t rely on them for warfare, but if necessary they are more than capable of acting as soldiers in large numbers…

Formica Fusca Soldier Ant (Ereptor)

The Fusca Soldier is the workhorse of the Fusca empire, its a capable ant with average health and an average attack strength. What it has over the worker is that its attack speed is higher, and when upgraded can deal much higher damage and attack faster while also enduring more hits than the worker can ever do.

It costs more food to plant but it pays for itself by being a staple soldier and capable asset to any army.

When controlled by the player in the Ereptor Hierarchy, Fusca Soldiers are the swarming melee soldiers. They are best used as front line combat ants, and meatshields for other weaker ants, or even a screening force for workers going about their buisness.

When upgraded they can either deal more damage or endure more damage, or even recover health while out of combat or retreat from combat when they lose a percentage of their overall health.

Formica Sanguinea – The Slave Maker Ants

Formica Sanguinea is a Non Player Controlled (NPC) Ant, and is only encountered during the Fusca Level in Challenge 1.2.

Formica Sanguinea are not really ‘dangerous’, per say, in that they dont have the numbers or strength to overpower the Black Ant soldiers 1 on 1, but they are dangerous to the colony because of their nasty ability and how they attack the colony.

They will attack in waves, charging into the nest in squads of increasing number as the colony progresses in time…

Sanguinea only have ‘1’ ant type that is encountered, a Slave Maker Soldier…

Slave Maker Soldier

Slave Maker Soldiers are after the colony not to exterminate per-say, but to steal larvae for their own colony. When they attack, half will engage the Colony Ants in battle, acting as a distraction while others rush and dash to the nursery tiles of the colony.

The Soldiers of the Sanguinea are on par with the Black Ant Soldiers in terms of health, damage, and speed. They are best engaged in standard Ant v Anything tactics, with numbers and unified with workers and soldiers combined.

If any of the Slave Makers that are going for the larvae manage to grab one, and get away, that tile will be put onto a cool down in which no larvae can be placed, and if the ant belonging to that tile is killed cannot be replaced until the cooldown is gone.

As stated before they do not pose a strong threat, but if Black Ant colony cannot get the numbers needed to push them back without suffering significant losses, the increasing raid strength will soon overwhelm the colony.

They cannot be encountered anywhere else and as such only are unique to level 1.2 of the black ant campaign…

Formica Rufa – Wood Ants with Range

Formica Rufa is the ‘2nd’ playable species that can be encountered and played as besides Fusca and not counting Ereptor.

Formica Rufa is special in that unlike Fusca, Rufa has more types of ants than just traditional Worker and Soldier, having 2 types of soldiers that are best employed ‘together’ to form a powerful army.

Formica Rufa Queen

She is like all other Queens, fat, sits there, lays eggs, if dies you lose gg

Formica Rufa – Worker

Formica Rufa Workers are like all other Workers, they do little damage, tehy have little health, and perform worker jobs, unlike Fusca workers though, they do not possess strong attacks, and are best employed as gatherers, performing worker jobs underground in the nest, or only… as early game frontline forces…

Formica Rufa – Melee Soldier

Formica Rufa soldiers, the melee variety, have extremely weak attacks, however they make up for this by having an extremely high health. This makes them ideal ants to act as ‘meat shields’, as they can endure a ton of damage compaerd to the rest of the Rufa force.

In particular they are best employed alongside their ranged counterpart… whether motor or rapid…

Formica Rufa – Ranged Motor (Ereptor)

Formica Rufa Motor Ants have low health, and are more expensive then their melee counterparts, but make up for this by using ‘ranged’ attacks instead of melee combat.

The Motor version use much slower attack speeds but have higher damage

when Upgraded the Motor Ants can deal splash damage as well.

Formica Rufa – Ranged Rapid (Ereptor)

Formica Rufa rapid ants have low health but use a much faster attacking, if low damage output.

The Rapid variety can be upgraded and when they are, they can deal a slowdown effect to targets they hit.


You can only pick ‘1’ of the rufa ranged ants to use, so when you decide you must choose either Motor or Rapid

Rapid deal faster damage in which the individual shots are weaker, while the Motor deals slower but ‘stronger’ shots.

Atta Cephalotes – The Ant Vegans

The newest Playable Ant species, and also the most complicated of them so far, Atta Cephalotes, also known as the Leaf Cutters, act very differently from every other ant species in the game.

As their name suggests, The Leaf Cutters get their food not from other insects, bits of meat from fish or even seeds… but from leaves.

This species harvests from plants by cutting up and bringing bits and pieces of leaves to its colony and drops them off, where they are picked up and taken to nursery tiles and become Fungus, which is what the colony uses for food.

As this species uses leaves for food, killing other insects will not offer them food of any kind.

As for the ants themselves in the species, Atta Cephalotes takes ant types to another level, and puts all of its Ants into various Castes, each with their own roles and purpose in the colony.

They are as Follows

Atta Cephalotes Queen

She’s Big, She’s Fat, She’s in internet terms EXTRA THICC… and just as useless outside of making eggs like the other queens. She dies, GG

Atta Cephalotes Minum Workers

The Minums are the lowest caste, as well as being the absolute tiniest ant of any species. These little girls do not leave the nest for any reason and nor can they be commanded at all. They cannot attack, they have little health, and perform the most important jobs that the other ants can’t do.

Minums go to leaf storage tiles and collects bits and pieces of leaves, and bring them to various nursery tiles to be cultivated into Fungus, which is used as food.

Minums also collect waste product that using fungus produces, and drops them off in ‘refuse’ chambers, where it sits and slowly disintegrates.

Minums also are the only ant that are unaffected by the Refuse Chambers negative effect on ants that are unfortunate enough to walk through them.

Atta Cephalotes Minor Caste

The Minors of the Leaf Cutters fullfill the role of other workers, they harvest, they plant nursery tiles, leaf storage tiles and plant waste refuse tiles on top of Highway tiles.

This caste of leaf cutter can harvest leaves like all leaf cutters, but because they are small they cannot carry heavy loads, and any leaf they carry will slow them down considerably.

They are poor attackers like all workers, but when in doubt, Numbers can make anything happen.

Atta Cephalotes Midae Caste

The Midae caste of the leaf cutters are considered a hybrid of soldier and worker, and are the workhorse of any Atta Cephalotes harvesting effort, capable of cutting larger chunks compared to their Minor sisters, without being slowed down.

In a pinch, the Midae Caste can easily fight off minor threats and in large enough numbers, can take on larger threats.

What they cannot do though, is take on major threats that are just too powerful to overcome even with numbers… That is where the Leaf Cutter Colony unleashes their final caste…

Atta Cephalotes Major Caste (Ereptor)

The Major Caste of the Leaf Cutters are the ultimate soldier, and are the backbone of any army in the Leaf Cutter Arsenal.

They are the largest, still dwarfed by the queen, but their powerful size and powerful jaws make them the ultimate force to be reckoned with on the field.

Though they are soldiers, the Major Caste can easily use their powerful muscles and slicing jaws to carve up leaves and carry chunks too large for even Midae to carry, back to their nest.

When the Ereptor Colony uses them, tehy can be upgraded to either draw in attacks toward them and away from other weaker ants, or can be used to ‘stun’ enemies smaller than them for a short duration.

Boasting high health and powerful but slow melee damage, they are the first into battle, and are often the last ones to fall…

Odontomachus Bauri – Trappers of the Undergrowth

Formica Odontomachus Bauri, more commonly known as Trap Jaw Ants, are only seen in the Leaf Cutter missions, in particular 3.1 while two warring leaf cutters compete

They typically roam the undergrowth in small groups of two or more, and rarely attack colonies and only engage leaf cutter ants if they happen to cross paths.

They only have ‘1’ Trap Jaw ant type that they reveal, their wandering Soldier class

Odontomachus Bauri Soldier

The soldier of the Trap Jaw ant sports a powerful set of pincers, designed to ensnare and slice into their targets.

While individually they are dangerous, against groups of ants these often solo hunters do not fair well against their enemies. What makes them unique and dangerous is their uncanny ability to ‘disengage’ from their enemy after taking a bit of damage.

Once their health drops below a certain point, they activate their leap ability, using their pincers to fling themselves away from their target. This allows them to temporarily retreat, but at the same time it regenerates some health before they reengage.

As time passes the Trap Jaw hunting parties will grow in number, making them more and more of a threat the longer you deal with them.

Eciton Burchellii – Roaming Ant Horde

Eciton Burchellii are by far among the ‘strongest’ ant species known thus far in the game

While the other ants possess a simliar type set to each other, workers and soldiers, Eciton Burchellii brings it to a whole new level.

Eciton Burchellii army ants are encountered only in 3.2 level of the leaf cutter ants, they make their presence and dangerous threat level known by simply ‘appearing’. They have a simliar Caste system to the Leaf Cutters, but unlike their vegan cousins, the Army Ant roams the undergrowth in search of prey.

Where they go the horde as a whole roams as well, setting up their nest and spreading out to consume and hunt down for whatever they can. Insects will flee from their paths, threats the player may fear will become trivial to them… and their power becomes known in their caste of ants…

Eciton Burchellii Scout Caste

The Scouts of the Army Ants are about the size of Minums for the Leaf Cutter, if not even smaller, and they have extremely low health and low attack.

By themselves they are no threat to even the lowliest caste in the Leaf Cutters, but they possess a unique power that makes them a priority threat to the player’s leaf cutter colony. If they encounter anything living that is a threat, or a potential target, they will retreat and lay down a trail to the location where they encountered the threat, bringing with them a small battalion of Army Ant Soldiers.

Eciton Burchellii Worker Caste

The Burchellii workers, like all workers, pose little offensive threat to their enemies, but in large numbers can become a massive threat.

They are seen walking along a trail guarded by the Majors of their colony, carrying food back to their nest or scavenging any poor creature foolish enough to wander into their hunting lines…

Eciton Burchellii Major Caste

Though only half the size of the Leaf Cutter Majors with less health compared to the massive warrior of the Vegan Ant colony, Eciton Burchellii Majors are just as recognizeable to anyone by their massive heads with powerful pincers.

Eciton Burchellii Majors may have less health compared to their leaf cutter major counterparts, but they compensate for this by having an even ‘stronger’ attack, and a unique ability. During combat if any Major engages in battle with others around it, after a while or after securing a kill, the Major will release a pheromone that encourages even faster attacks by allied ants around it… Making the Army Ant Major, the greatest threat to any ant colony they come across.

Unless overwhelmed quickly, a single Major surrounded by dozens of workers can overpower even mighty ant armies…

Player Ant Strategies

When playing as any of the playable ant colonies, Ereptor, Fusca, Rufa, or Atta Cephalotes, one must have an overall strategy.

We wont cover the Ereptor Ants because that is player choice, and as such the only thing that stops you from becoming a dominating force is your lack of upgrades and ant species…

Fusca, Rufa, Atta Cephalotes Strategies

Formica Fusca

Formica Fusca, the Black Ants, are the most basic of basic ant colony species in the Undergrowth, only possessing 2 types of ants, workers and soldiers.

Their playstyle is as simple as it is basic, build up your workers and soldiers, and use the only thing they have to their advantage… Superior Numbers…

As they dont need overcomplicated strategies or plans, they have to simply build up a large army and swarm their enemies using superior numbers and overwhelming force.

Formica Rufa

Formica Rufa, or the Wood Ants, can use the same superior numbers playbook as the Black Ants, but combine it with army composition.

While one common plan is to use the cheap workers as simple fodder for your ranged ants, which deal the real damage of the Rufa Colony, they do not have the health needed to endure a sustained attack for long. The only way this works is if you invest ‘heavily’ into workers, and can keep up the sustained losses they will be incurring.

The real meatshields are in their melee soldiers, and should act as the frontline of your army.

For ranged Rufa’s motor or rapid fire ants each deliver damage in their own ways, either dealing lots of smaller hits or massive damage in slow volleys. Upgraded they can also deal either slowing venom damage or splash damage.

One way to maxamize their damage output is to have just slightly ‘more’ ranged rufa ants compared to melee rufa ants.

Atta Cephalotes

Atta Cephalotes possess a unique if somewhat ‘slower’ way of producing food.

Their caste system also makes it so that most of the times there is always someone harvesting leaves, but the Major’s time should be spent defending and only harvesting if the threat to the leaf cutter line is minimal.

Because of how Expensive Majors are (150 just to ‘plant’ a tile, and 15 to hatch a larvae) they are often not the ant type you will be buying a ‘lot’ of.

The Minors being cheapest, 20 / 2, will be doing a lot of the harvesting, alongside their larger cousins the Midae, 80 / 6.

When it comes to warfare, in particular the colony v colony warfare, its all hands on deck as because of how slow it is for fungus to grow and how expensive Majors and Midae are to plant, you will need to use swarming tactics to win, using every ant in the harvesting line to attack unless your income is high enough to support massive numbers of Midae alongside Majors.

While the others can rely on ‘quantity’ to win most battles, the Leaf Cutters should focus on ‘Quality’, upgrading their majors first and Midae second, as for every Major you lose its another 15 food to hatch another, and if the harvesting line is constantly under attack by stray bugs you’ll be spending 2 for every worker, 5-6 for every Midae… which piles up fast and can deplete your fungus reserves quickly

Formica Sanguinea, Odontomachus Bauri, Eciton Burchellii

As you cannot play as these ant colonies, the only thing you can do is play against them.

Fighting against these 3 ant species has their own challenges

Formica Sanguinea

fighting against Formica Sanguinea is like fighting against a pair of pick pocket thieves.

You need to have 1 hand hold off one guy while the other fights off against the other guy.

The strategy when fighting against this ant species is simply to have higher numbers, and use everyone to attack them, while one group holds off their attackers, another one holds off and attacks those raiding the larvae nests.

They aren’t that dangerous when numbers are on your side.

Odontomachus Bauri

Odontomachus Bauri trapjaw ants are not like the other ant colony enemies, they do not engage in colony warfare at this time in the game. All they do is roam the undergrowth in small groups of Ants.

Soldiers will leave their nests to scour around for food.

Their unique method of fighting, attacking and then leaping away to recover health means that to defeat them you need to consistantly engage them. If you can outnumber them while keeping them under constant attack, their ability to regenerate health will be minimal.

Eciton Burchellii

Eciton Burchellii are by far the greatest threat your colony will face, their numbers, their powerful attack, their hoard mentality, and the fact that if even ‘1’ scout escapes it will bring in a battalion of Army Ants, makes them a threat to be respected.

The first thing you need to do is prepare for them, do not engage the Army Ants until you have a sufficient number of ants, and when you battle them you have to commit to throwing them back using ‘everything’ you have, Leaf Cutter Minors, Midae, and Majors together to fight them as using only the Majors will result in them being overwhelmed by the horde.

Because in 3.2 the horde of Army Ants will continue to assault a path until they’ve taken sufficient losses, you will need to keep the pressure up on them. To that end numbers alone wont be enough, you will need quality on your side, and one way to have that is to upgrade your Majors to counter ‘their’ Majors. Remember that Major Ants are the most expensive to rebuild of all the Ants in the Leaf Cutter arsenal, so investing into an upgraded batch of Majors is going to be a better investment then having a ton of Majors that will quickly drain your food reserves in a sustained fight against the Army Ants.

Speaking of Majors, Eciton Burchellii majors are by far the strongest and scariest of all the ants you’ve faced. They improve the attacking power of allies, they hit hard, and they never stop coming…

Their one weakness is low health, so if you can overwhelm them quickly their ability to boost damage will be lost.

When pushing the swarm back, be careful when you engage, as not only Workers but Majors will travel along the path together, so if you are fighting on one side you can find yourself in a pincer between two waves of Army Ants.


This is the ending of Empires Of the Undergrowth All Species of Ants guide. I hope it will help you. If there is wrong or you have suggestions, please let’s know and comment us. Have fun.

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Written by Imperatordavius

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